Pyrularia pubera

ClassificationDicots
FamilySantalaceae
Family (Traditional)Santalaceae
GenusPyrularia
SpeciesPyrularia pubera Michx.
Common NameBuffalo nut
Growth Habitshrub
Plant NotesLeaf Persistance: Deciduous; Habit: shrub; Woody/Herbaceous: woody; Terrestrial/Aquatic: Terrestrial;
Leaf Arrangement: Alternate; Simple/Compound: Simple; Leaf Complexity: 0; Leaf Type: Broadleaf; Leaf Veination: Pinnate; Petiole Type: Petiolate; Leaf/Leaflet Shapes: Elliptic;Lanceolate;Oblanceolate;Oblong;Obovate; Leaf/Leaflet Margins: Entire;
Hypanthium: No; Perianth Absent: No; Fruit Color: ;
*Average C-Value: 8.0
C value 8
StatusNative; AGCP (FACU), EMP (UPL) (NWPL); TN[SNR] KY[S5] (State Rank); G5 (Global Rank)
Specimen(s) View and Download Specimen Details »

** Not applicable or data not available.

Species Distribution Map

This species has been reported in the counties highlighted in the interactive distribution map below. Click on a county to display the its name. Click the question mark icon () to view additional information about the ecoregions shown. In the map legend, there are links to toggle the visibility of the ecoregions and/or their labels. In addition, clicking on an ecoregion name will highlight that region on the map.

  • State Boundaries
  • Counties
  • Counties - Species Present
Ecoregions
  • Southeastern Plains
    • Blackland Prarie
  • Blue Ridge Mountains
  • Ridge and Valley
  • Cumberland Plateau
  • Cumberland Mountains
  • Allegheny Plateau
  • Interior Plateau
    • Bluegrass Region
    • Nashville Basin
    • Pennyroyal Karst Plain
    • Western Highland Rim
    • Eastern Highland Rim

Synonyms

No synonyms exist for this species.

Specimens and Distribution

This species has been reported in the following counties by the herbaria listed. An overview of the individual specimens are provided in the table that follows. Click on the accession number to view details; click on column headers to sort.

Counties included on distribution map: Bell (KY), Breathitt (KY), Clay (KY), Elliott (KY), Harlan (KY), Jackson (KY), Johnson (KY), Knott (KY), Laurel (KY), Lee (KY), Leslie (KY), Letcher (KY), McCreary (KY), Magoffin (KY), Menifee (KY), Owsley (KY), Perry (KY), Pike (KY), Powell (KY), Rockcastle (KY), Whitley (KY), Wolfe (KY), Anderson (TN), Blount (TN), Campbell (TN), Carter (TN), Claiborne (TN), Cocke (TN), Cumberland (TN), Fentress (TN), Greene (TN), Grundy (TN), Hamilton (TN), Hancock (TN), Jefferson (TN), Johnson (TN), McMinn (TN), Marion (TN), Monroe (TN), Morgan (TN), Polk (TN), Rhea (TN), Roane (TN), Scott (TN), Sevier (TN), Sullivan (TN), Unicoi (TN), Washington (TN)

Herbaria represented by specimen data listed below: APSC, BEREA, EKY, ETSU, HTTU, KNK, MTSU, TENN, UCHT

Range of years during which specimens were collected: 1948 - 2017

Accession No. (click for full details) County State Coll. Date Collector &
Collection No.
Herbarium &
Herbarium Name Used
31234100644663 Harlan Kentucky 22 May 1993 Bryce D. Fields
279
EKY
Pyrularia pubera
31234100644978 Pike Kentucky 15 May 1965 Elizabeth M. Browne
9994
EKY
Pyrularia pubera
31234100644648 Harlan Kentucky 22 May 1993 Bryce D. Fields
269
EKY
Pyrularia pubera
31234100644804 Leslie Kentucky 08 May 2006 Timothy J. Weckman
10159
EKY
Pyrularia pubera
N/A Harlan Kentucky 09 Aug 1999 Timothy J. Weckman
5344
EKY
Pyrularia pubera
31234100644853 McCreary Kentucky 20 Apr 1979 Richard H. Hannan
1345
EKY
Pyrularia pubera
31234100645025 Wolfe Kentucky 10 Oct 1964 Elizabeth M. Browne
9529.5
EKY
Pyrularia pubera
31234100644937 Perry Kentucky 13 Jun 1979 Richard H. Hannan
2267
EKY
Pyrularia pubera
31234100644911 Owsley Kentucky 26 May 1996 Timothy J. Weckman
2625
EKY
Pyrularia pubera
31234100644846 Magoffin Kentucky 21 Jun 1979 Richard H. Hannan
2481
EKY
Pyrularia pubera
31234100644606 Elliott Kentucky 15 Oct 2012 M. Greene
EKY
Pyrularia pubera
31234100644564 Breathitt Kentucky 05 Aug 1963 Elizabeth M. Browne
7823
EKY
Pyrularia pubera
31234100859113 Wolfe Kentucky 24 Jul 1972 Jerry Buskin
1394
EKY
Pyrularia pubera
31234100644671 Harlan Kentucky 30 Jul 2004 Amy V. McIntosh
Jacob Stewart
EKY
Pyrularia pubera
31234100644572 Breathitt Kentucky 27 Jun 1979 Richard H. Hannan
2686
EKY
Pyrularia pubera
31234100644879 McCreary Kentucky 18 May 1996 Timothy J. Weckman
2583
EKY
Pyrularia pubera
31234100645017 Wolfe Kentucky 13 Jul 1979 Richard H. Hannan
2988
EKY
Pyrularia pubera
31234100644622 Harlan Kentucky 27 Sep 1978 Richard H. Hannan
01-013-045
EKY
Pyrularia pubera
31234100644945 Perry Kentucky 01 Jan 1995 Morris Elsea
EKY
Pyrularia pubera
31234100644556 Bell Kentucky 14 May 1974 J. Stuart Lassetter
2059
EKY
Pyrularia pubera
31234100644952 Pike Kentucky 21 May 1964 Elizabeth M. Browne
8450
EKY
Pyrularia pubera
31234100644796 Lee Kentucky 28 May 1993 Timothy J. Weckman
249
EKY
Pyrularia pubera
31234100746054 McCreary Kentucky 02 Sep 2008 Tara R. Littlefield
2008-051
EKY
Pyrularia pubera
31234100644903 McCreary Kentucky 14 Oct 1990 Douglas Stephens
48
EKY
Pyrularia pubera
31234100644689 Harlan Kentucky 09 Aug 1999 Timothy J. Weckman & Judith E. Weckman
5344
EKY
Pyrularia pubera
31234100644754 Knott Kentucky 07 Jul 1996 Timothy J. Weckman
3000
EKY
Pyrularia pubera
31234100645140 Morgan Tennessee 09 Aug 1990 Ronald L. Jones
6594
EKY
Pyrularia pubera
31234100644697 Harlan Kentucky 01 Oct 1948 Mary E. Wharton
6303
EKY
Pyrularia pubera
31234100644820 Letcher Kentucky 18 May 1978 Jeffrey D. Sole
64
EKY
Pyrularia pubera
31234100644986 Powell Kentucky 19 Jul 1991 Carol J. Way
136
EKY
Pyrularia pubera
31234100852860 Powell Kentucky 14 Jul 1985 P. Froelich
154
EKY
Pyrularia pubera
31234100645157 Scott Tennessee 13 May 1999 Joey Shaw
61
EKY
Pyrularia pubera
31234100778735 Wolfe Kentucky 09 May 2017 Thomas McFadden
470
EKY
Pyrularia pubera
31234100644580 Clay Kentucky 02 Jul 1980 Richard H. Hannan
4355
EKY
Pyrularia pubera
31234100644721 Harlan Kentucky 23 Aug 1996 R. L. Jones
7966
EKY
Pyrularia pubera
31234100644994 Rockcastle Kentucky 30 May 1979 Richard H. Hannan
1987
EKY
Pyrularia pubera
31234100645009 Rockcastle Kentucky 15 May 1995 Timothy J. Weckman
1413
EKY
Pyrularia pubera
31234100852878 Wolfe Kentucky 13 Jul 1985 Ross C. Clark
20980
EKY
Pyrularia pubera
31234100644788 Lee Kentucky 27 Apr 1996 Danny Husband
22
EKY
Pyrularia pubera
31234100644762 Lee Kentucky 28 May 1993 Timothy J. Weckman
282
EKY
Pyrularia pubera
31234100644770 Lee Kentucky 30 Jun 1996 Timothy J. Weckman
2929
EKY
Pyrularia pubera
31234100644747 Johnson Kentucky 05 Jun 1979 Richard H. Hannan
2073
EKY
Pyrularia pubera
31234100644739 Jackson Kentucky 26 Aug 1997 Ronald L. Jones
8209
EKY
Pyrularia pubera
31234100644895 McCreary Kentucky 28 Apr 1984 Ronald L. Jones
4234
EKY
Pyrularia pubera
31234100644838 Letcher Kentucky 16 May 1978 Jeffrey D. Sole
17
EKY
Pyrularia pubera
31234100644861 McCreary Kentucky 24 Jun 1972 Elizabeth M. Browne
72E10.23
EKY
Pyrularia pubera
31234100644812 Leslie Kentucky 23 May 1979 Richard H. Hannan
1911
EKY
Pyrularia pubera
31234100644929 Owsley Kentucky 28 Jun 1979 Richard H. Hannan
2746
EKY
Pyrularia pubera
31234100644705 Harlan Kentucky 06 Aug 1998 Ronald L. Jones
8574
EKY
Pyrularia pubera
31234100644960 Pike Kentucky 24 Jun 1965 Elizabeth M. Browne
10422.7
EKY
Pyrularia pubera
ETSU007755 Unicoi Tennessee 06 Aug 2009 Frosty Levy
ETSU
Pyrularia pubera
31973000018762 Harlan Kentucky
KNK
Pyrularia pubera
31973000018763 Cumberland Tennessee
KNK
Pyrularia pubera
31973000018757 McCreary Kentucky
KNK
Pyrularia pubera
31973000018758 Harlan Kentucky
KNK
Pyrularia pubera
31973000018756 Sevier Tennessee
KNK
Pyrularia pubera
31234100644630 Harlan Kentucky 22 May 1993 Bryce D. Fields
251
EKY
Pyrularia pubera
31234100644614 Elliott Kentucky 07 Jun 1979 Richard H. Hannan
2168
EKY
Pyrularia pubera
31234100644655 Harlan Kentucky 20 May 1962 Elizabeth M. Browne
5351
EKY
Pyrularia pubera
31234100644598 Elliott Kentucky 06 Apr 2013 M. Greene
EKY
Pyrularia pubera
BEREA021890 Knott Kentucky
BEREA
Pyrularia pubera
N/A Greene Tennessee 12 May 2009 Frosty Levy
ETSU
Pyrularia pubera
31973000018755 McCreary Kentucky
KNK
Pyrularia pubera
31973000018761 McCreary Kentucky
KNK
Pyrularia pubera
31973000018760 Harlan Kentucky
KNK
Pyrularia pubera
31973000018759 Letcher Kentucky
KNK
Pyrularia pubera
TENN-V-0085609 McMinn Tennessee
TENN
Pyrularia pubera
TENN-V-0085582 Claiborne Tennessee
TENN
Pyrularia pubera
TENN-V-0085573 Blount Tennessee
TENN
Pyrularia pubera
TENN-V-0085618 Polk Tennessee
TENN
Pyrularia pubera
TENN-V-0085591 Cumberland Tennessee
TENN
Pyrularia pubera
TENN-V-0085585 Claiborne Tennessee
TENN
Pyrularia pubera
TENN-V-0085622 Rhea Tennessee
TENN
Pyrularia pubera
TENN-V-0085612 Morgan Tennessee
TENN
Pyrularia pubera
TENN-V-0085655 Pike Kentucky
TENN
Pyrularia pubera
TENN-V-0085620 Polk Tennessee
TENN
Pyrularia pubera
TENN-V-0085633 Sevier Tennessee
TENN
Pyrularia pubera
TENN-V-0085574 Blount Tennessee
TENN
Pyrularia pubera
TENN-V-0085595 Fentress Tennessee
TENN
Pyrularia pubera
TENN-V-0085652 Breathitt Kentucky
TENN
Pyrularia pubera
TENN-V-0085650 Bell Kentucky
TENN
Pyrularia pubera
TENN-V-0085572 Blount Tennessee
TENN
Pyrularia pubera
TENN-V-0085602 Jefferson Tennessee
TENN
Pyrularia pubera
TENN-V-0085578 Campbell Tennessee
TENN
Pyrularia pubera
TENN-V-0085571 Blount Tennessee
TENN
Pyrularia pubera
TENN-V-0085638 Unicoi Tennessee
TENN
Pyrularia pubera
TENN-V-0085594 Fentress Tennessee
TENN
Pyrularia pubera
TENN-V-0085610 Monroe Tennessee
TENN
Pyrularia pubera
TENN-V-0085615 Morgan Tennessee
TENN
Pyrularia pubera
TENN-V-0085587 Cumberland Tennessee
TENN
Pyrularia pubera
TENN-V-0085632 Sevier Tennessee
TENN
Pyrularia pubera
TENN-V-0085569 Anderson Tennessee
TENN
Pyrularia pubera
TENN-V-0085637 Unicoi Tennessee
TENN
Pyrularia pubera
TENN-V-0085575 Blount Tennessee
TENN
Pyrularia pubera
TENN-V-0085588 Cumberland Tennessee
TENN
Pyrularia pubera
TENN-V-0085656 Wolfe Kentucky
TENN
Pyrularia pubera
TENN-V-0085640 Washington Tennessee
TENN
Pyrularia pubera
TENN-V-0085611 Monroe Tennessee
TENN
Pyrularia pubera
TENN-V-0085625 Roane Tennessee
TENN
Pyrularia pubera
TENN-V-0085624 Rhea Tennessee
TENN
Pyrularia pubera
TENN-V-0085651 Bell Kentucky
TENN
Pyrularia pubera
31234100644713 Harlan Kentucky 19 Aug 1998 Ronald L. Jones
8813
EKY
Pyrularia pubera
31234100644887 McCreary Kentucky 28 Apr 1984 Ronald L. Jones
4234
EKY
Pyrularia pubera
TENN-V-0085586 Cocke Tennessee
TENN
Pyrularia pubera
TENN-V-0085576 Campbell Tennessee
TENN
Pyrularia pubera
TENN-V-0085570 Anderson Tennessee
TENN
Pyrularia pubera
TENN-V-0085649 Bell Kentucky
TENN
Pyrularia pubera
TENN-V-0085653 Harlan Kentucky
TENN
Pyrularia pubera
TENN-V-0085584 Claiborne Tennessee
TENN
Pyrularia pubera
TENN-V-0085623 Rhea Tennessee
TENN
Pyrularia pubera
TENN-V-0085616 Morgan Tennessee
TENN
Pyrularia pubera
TENN-V-0085627 Scott Tennessee
TENN
Pyrularia pubera
TENN-V-0085577 Campbell Tennessee
TENN
Pyrularia pubera
TENN-V-0085613 Morgan Tennessee
TENN
Pyrularia pubera
TENN-V-0085628 Scott Tennessee
TENN
Pyrularia pubera
TENN-V-0085603 Jefferson Tennessee
TENN
Pyrularia pubera
TENN-V-0085590 Cumberland Tennessee
TENN
Pyrularia pubera
TENN-V-0085626 Scott Tennessee
TENN
Pyrularia pubera
TENN-V-0085631 Sevier Tennessee
TENN
Pyrularia pubera
TENN-V-0085601 Hancock Tennessee
TENN
Pyrularia pubera
TENN-V-0085598 Greene Tennessee
TENN
Pyrularia pubera
TENN-V-0085606 Johnson Tennessee
TENN
Pyrularia pubera
TENN-V-0085617 Polk Tennessee
TENN
Pyrularia pubera
TENN-V-0085629 Sevier Tennessee
TENN
Pyrularia pubera
TENN-V-0085580 Carter Tennessee
TENN
Pyrularia pubera
TENN-V-0085621 Polk Tennessee
TENN
Pyrularia pubera
TENN-V-0085605 Johnson Tennessee
TENN
Pyrularia pubera
TENN-V-0085630 Sevier Tennessee
TENN
Pyrularia pubera
TENN-V-0085635 Unicoi Tennessee
TENN
Pyrularia pubera
TENN-V-0085639 Washington Tennessee
TENN
Pyrularia pubera
TENN-V-0085589 Cumberland Tennessee
TENN
Pyrularia pubera
TENN-V-0085583 Claiborne Tennessee
TENN
Pyrularia pubera
TENN-V-0085604 Johnson Tennessee
TENN
Pyrularia pubera
TENN-V-0085592 Cumberland Tennessee
TENN
Pyrularia pubera
TENN-V-0085654 McCreary Kentucky
TENN
Pyrularia pubera
TENN-V-0085596 Fentress Tennessee
TENN
Pyrularia pubera
TENN-V-0085593 Cumberland Tennessee
TENN
Pyrularia pubera
TENN-V-0085599 Grundy Tennessee
TENN
Pyrularia pubera
TENN-V-0085597 Greene Tennessee
TENN
Pyrularia pubera
TENN-V-0085619 Polk Tennessee
TENN
Pyrularia pubera
TENN-V-0085607 Johnson Tennessee
TENN
Pyrularia pubera
TENN-V-0085634 Sullivan Tennessee
TENN
Pyrularia pubera
TENN-V-0085608 Marion Tennessee
TENN
Pyrularia pubera
TENN-V-0085579 Campbell Tennessee
TENN
Pyrularia pubera
UCHT010494 Hamilton Tennessee
UCHT
Pyrularia pubera
UCHT010481 Cumberland Tennessee
UCHT
Pyrularia pubera
UCHT010487 Polk Tennessee
UCHT
Pyrularia pubera
TENN-V-0085581 Carter Tennessee
TENN
Pyrularia pubera
TENN-V-0085614 Morgan Tennessee
TENN
Pyrularia pubera
TENN-V-0085636 Unicoi Tennessee
TENN
Pyrularia pubera
TENN-V-0085648 Bell Kentucky
TENN
Pyrularia pubera
UCHT010484 Polk Tennessee
UCHT
Pyrularia pubera
UCHT010490 Polk Tennessee
UCHT
Pyrularia pubera
UCHT010482 Cumberland Tennessee
UCHT
Pyrularia pubera
UCHT010491 Tennessee
UCHT
Pyrularia pubera
APSC0085636 McCreary Kentucky
APSC
Pyrularia pubera
APSC0003569 Blount Tennessee 26 Apr 1968 EW Chester with Ken Rogers
2002
APSC
Pyrularia pubera
APSC0119198 Pike Kentucky
APSC
Pyrularia pubera
APSC0103984 McCreary Kentucky
APSC
Pyrularia pubera
APSC0018342 Polk Tennessee 29 Sep 2010 Joey Shaw, Dwayne Estes, Eugene Wofford, Sunny Hart, Emily Blyveis, Aaron Floden, and Rob Shepard
APSC
Pyrularia pubera
APSC0119197 Harlan Kentucky
APSC
Pyrularia pubera
APSC0080041 Knott Kentucky 13 Nov 1986
APSC
Pyrularia pubera
UCHT010486 Hamilton Tennessee
UCHT
Pyrularia pubera
UCHT010489 Tennessee
UCHT
Pyrularia pubera
UCHT010493 Sevier Tennessee
UCHT
Pyrularia pubera
UCHT010483 Polk Tennessee
UCHT
Pyrularia pubera
APSC0117972 Bell Kentucky
APSC
Pyrularia pubera
APSC0057303 Morgan Tennessee
APSC
Pyrularia pubera
APSC0085620 Pike Kentucky
APSC
Pyrularia pubera
APSC0103982 Harlan Kentucky
APSC
Pyrularia pubera
APSC0103987 Harlan Kentucky
APSC
Pyrularia pubera
APSC0111163 Lee Kentucky
APSC
Pyrularia pubera
APSC0119195 Johnson Kentucky
APSC
Pyrularia pubera
APSC0119200 Wolfe Kentucky
APSC
Pyrularia pubera
APSC0103986 Whitley Kentucky
APSC
Pyrularia pubera
APSC0055574 Cumberland Tennessee
APSC
Pyrularia pubera
APSC0119202 Harlan Kentucky
APSC
Pyrularia pubera
APSC0085632 Harlan Kentucky 02 Aug 1984
APSC
Pyrularia pubera
APSC0085634 Menifee Kentucky
APSC
Pyrularia pubera
APSC0094684 Whitley Kentucky
APSC
Pyrularia pubera
APSC0103146 Bell Kentucky
APSC
Pyrularia pubera
APSC0103985 Letcher Kentucky
APSC
Pyrularia pubera
APSC0085635 Menifee Kentucky
APSC
Pyrularia pubera
APSC0119196 Harlan Kentucky
APSC
Pyrularia pubera
APSC0119201 Wolfe Kentucky
APSC
Pyrularia pubera
APSC0003568 Blount Tennessee 01 May 1964 Wayne Chester with Dale Thomas
134
APSC
Pyrularia pubera
APSC0117973 Wolfe Kentucky
APSC
Pyrularia pubera
APSC0119206 Wolfe Kentucky
APSC
Pyrularia pubera
APSC0117974 McCreary Kentucky
APSC
Pyrularia pubera
APSC0119203 Letcher Kentucky
APSC
Pyrularia pubera
APSC0119204 McCreary Kentucky
APSC
Pyrularia pubera
APSC0045058 Cumberland Tennessee
APSC
Pyrularia pubera
APSC0003567 Cumberland Tennessee 26 Apr 1968 Wayne Chester
2006
APSC
Pyrularia pubera
APSC0086010 Bell Kentucky
APSC
Pyrularia pubera
APSC0085633 Laurel Kentucky
APSC
Pyrularia pubera
APSC0119199 Wolfe Kentucky
APSC
Pyrularia pubera
APSC0003570 Scott Tennessee 07 May 1999 Joey Shaw with Chris Fleming
36
APSC
Pyrularia pubera
APSC0080042 Knott Kentucky 13 Nov 1986
APSC
Pyrularia pubera
APSC0003571 Fentress Tennessee 28 Jul 1999 Joey Shaw with Chris Fleming
359
APSC
Pyrularia pubera
HTTU015873 Scott Tennessee
HTTU
Pyrularia pubera
HTTU029395 Scott Tennessee
HTTU
Pyrularia pubera
HTTU015867 Scott Tennessee
HTTU
Pyrularia pubera
HTTU015870 Fentress Tennessee
HTTU
Pyrularia pubera
HTTU029396 Scott Tennessee
HTTU
Pyrularia pubera
HTTU015872 Scott Tennessee
HTTU
Pyrularia pubera
HTTU015871 Scott Tennessee
HTTU
Pyrularia pubera
MTSU015846 Morgan Tennessee 23 Apr 1988 Duane Douglas Yeary
MTSU
Pyrularia pubera
UCHT030730 Tennessee
UCHT
Pyrularia pubera
UCHT036295 Polk Tennessee
UCHT
Pyrularia pubera
HTTU015869 Fentress Tennessee
HTTU
Pyrularia pubera
HTTU015868 Fentress Tennessee
HTTU
Pyrularia pubera
MTSU015847 Morgan Tennessee Duane Douglas Yeary
MTSU
Pyrularia pubera
UCHT036296 Polk Tennessee
UCHT
Pyrularia pubera
UCHT030729 Tennessee
UCHT
Pyrularia pubera
UCHT030732 Tennessee
UCHT
Pyrularia pubera
UCHT030733 Tennessee
UCHT
Pyrularia pubera
UCHT030727 Tennessee
UCHT
Pyrularia pubera
UCHT030734 Tennessee
UCHT
Pyrularia pubera
UCHT030731 Tennessee
UCHT
Pyrularia pubera
UCHT035742 Campbell Tennessee
UCHT
Pyrularia pubera
UCHT030728 Tennessee
UCHT
Pyrularia pubera
UCHT030735 Polk Tennessee
UCHT
Pyrularia pubera
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Plant Photos

Pyrularia pubera - Ashley B. Morris
Pyrularia pubera - Ashley B. Morris
Pyrularia pubera - Dwayne Estes
Pyrularia pubera - Dwayne Estes
Pyrularia pubera - Joey Shaw
Pyrularia pubera - Joey Shaw
Pyrularia pubera - Joey Shaw
Pyrularia pubera - Joey Shaw
Pyrularia pubera - Joey Shaw
Pyrularia pubera - Joey Shaw
Pyrularia pubera - Joey Shaw
Pyrularia pubera - Joey Shaw
Pyrularia pubera - Joey Shaw
Pyrularia pubera - Joey Shaw
Pyrularia pubera - Joey Shaw
Pyrularia pubera - Joey Shaw
Pyrularia pubera - Milo Pyne
Pyrularia pubera - Milo Pyne
Pyrularia pubera - Milo Pyne
Pyrularia pubera - Milo Pyne