Carduus nutans

ClassificationDicots
FamilyAsteraceae
Family (Traditional)Asteraceae
GenusCarduus
Species:Carduus nutans L.
Common NameMusk Or Nodding Thistle; Nodding Plumeless Thistle
Growth Habitforb
Plant Notes**
CharacteristicsArmaments: Yes; Milky Sap: No; Photosynthetic: Yes; Stipules: No; Woody Herbaceous: Herbaceous; Invasive Rank: KY-EPPC Severe Threat; Natural Area Presence: GILES; Natural Area Presence: GRSM; Natural Area Presence: LC; Natural Area Presence: LGS; Natural Area Presence: LULA; Natural Area Presence: PK; Duration: Annual; General Ecology: Terrestrial; Growth Habit: forb; Moisture Requirements: Sub-xeric; Moisture Requirements: Dry-mesic; Moisture Requirements: Mesic; Region of Origin: Asia; Region of Origin: Europe; Reproductive Phenology: May; Reproductive Phenology: June; Reproductive Phenology: July; Reproductive Phenology: August; Reproductive Phenology: September; Leaf Arrangement: alternate; Leaf Arrangement: basal; Leaf Complexity: simple; Leaf Dissection: 0.5; Leaf Fragrance: No; Leaf Persistance: Deciduous; Leaf Petiole Type: petiolate; Leaf Petiole Type: sessile; Leaf Type: broadleaf; Leaf Type: thistle-like; Leaf Venation: pinnate; Leaflet Margins: dentate; Leaflet Margins: entire; Leaflet Margins: lobed; Leaflet Number: 0; Anther Type: Basifixed; Base Flower Color: Purple; Carpel Number/Pistil: 1; Fruit Complexity: Simple; Fruit Dehiscence: No; Fruit Type: Cypsela; Fruit Wet/Dry: Dry; Hypanthium Present: No; Inflorescence Position: axillary; Inflorescence Position: terminal; Inflorescence Types: corymb/cyme/umbel; Inflorescence Types: head; Inflorescence Types: solitary; Ovary Position: Inferior; Overall Flower Symmetry: Actinomorphic; Ovules per Carpel: 1; Perianth Present: Yes; Petal Fusion: Yes; Petal Number: 5; Pistil Complexity: Simple; Pistil Number: 1; Placentation Type: Basal; Sepal Fusion: N/A; Sepal Number: N/A; Sexuality: monoclinous; Sporangium Arrangement: N/A; Stamen Number: 5; Tepal Number: N/A
C value N/A
StatusNot Native; Invasive; AGCP (UPL), EMP (UPL) (NWPL); TN[SNA] KY[SNA] (State Rank); GNR (Global Rank)
Specimen(s) View and Download Specimen Details »

** Not applicable or data not available.

Species Distribution Map

This species has been reported in the counties highlighted in the interactive distribution map below. Click on a county to display the its name. Click the question mark icon () to view additional information about the ecoregions shown. In the map legend, there are links to toggle the visibility of the ecoregions and/or their labels. In addition, clicking on an ecoregion name will highlight that region on the map.

  • State Boundaries
  • Counties
  • Counties - Species Present
Ecoregions
  • Southeastern Plains
    • Blackland Prarie
  • Blue Ridge Mountains
  • Ridge and Valley
  • Cumberland Plateau
  • Cumberland Mountains
  • Allegheny Plateau
  • Interior Plateau
    • Bluegrass Region
    • Nashville Basin
    • Pennyroyal Karst Plain
    • Western Highland Rim
    • Eastern Highland Rim

Synonyms

SynonymFull Citation
Carduus macrocephalus Carduus macrocephalus Coss.
Carduus macrolepis Carduus macrolepis Peterm.
Carduus nutans subsp. leiophyllus Carduus nutans subsp. leiophyllus
Carduus nutans subsp. macrocephalus Carduus nutans subsp. macrocephalus
Carduus nutans subsp. macrolepis Carduus nutans subsp. macrolepis
Carduus nutans subsp. nutans Carduus nutans subsp. nutans
Carduus nutans var. leiophyllus Carduus nutans var. leiophyllus
Carduus nutans var. macrocephalus Carduus nutans var. macrocephalus (Desf.) B.Boivin
Carduus nutans var. vestitus Carduus nutans var. vestitus (Hallier) B.Boivin
Carduus thoermeri Carduus thoermeri Weinm.

Specimens and Distribution

This species has been reported in the following counties by the herbaria listed. An overview of the individual specimens are provided in the table that follows. Click on the accession number to view details; click on column headers to sort.

Counties included on distribution map: Anderson (KY), Barren (KY), Campbell (KY), Carroll (KY), Christian (KY), Clark (KY), Clinton (KY), Estill (KY), Fayette (KY), Fleming (KY), Franklin (KY), Gallatin (KY), Garrard (KY), Hickman (KY), Jefferson (KY), Kenton (KY), Larue (KY), Logan (KY), Madison (KY), Marion (KY), Marshall (KY), Mercer (KY), Monroe (KY), Montgomery (KY), Muhlenberg (KY), Nelson (KY), Oldham (KY), Powell (KY), Rockcastle (KY), Rowan (KY), Scott (KY), Shelby (KY), Trigg (KY), Trimble (KY), Warren (KY), Benton (TN), Bledsoe (TN), Campbell (TN), Cannon (TN), Carroll (TN), Coffee (TN), Davidson (TN), DeKalb (TN), Dyer (TN), Fentress (TN), Franklin (TN), Giles (TN), Greene (TN), Grundy (TN), Hamblen (TN), Hamilton (TN), Haywood (TN), Henderson (TN), Houston (TN), Jackson (TN), Jefferson (TN), Knox (TN), Loudon (TN), Macon (TN), Marion (TN), Maury (TN), Montgomery (TN), Putnam (TN), Rutherford (TN), Sequatchie (TN), Sevier (TN), Shelby (TN), Smith (TN), Stewart (TN), Sullivan (TN), Sumner (TN), Unicoi (TN), Van Buren (TN), Washington (TN), White (TN), Williamson (TN), Wilson (TN)

Herbaria represented by specimen data listed below: APSC, BEREA, BOON, EKY, ETSU, HTTU, KNK, MDKY, MEM, MTSU, SWMT, TENN, UCHT, UOS

Range of years during which specimens were collected: 1956 - 2016

Accession No. (click for full details) County State Coll. Date Collector &
Collection No.
Herbarium &
Herbarium Name Used
N/A Madison Kentucky 16 Jun 1993 Gary W. Libby
422
EKY
Carduus nutans
31234100244902 Powell Kentucky 05 May 2001 Timothy J. Weckman
6305
EKY
Carduus nutans
31234100244738 Estill Kentucky 10 Jun 1987 Richard G. Guetig
176
EKY
Carduus nutans
31234100244811 Madison Kentucky 12 Jun 1993 Susan Wind
89
EKY
Carduus nutans
BEREA002865 Monroe Kentucky
BEREA
Carduus nutans
BEREA002862 Madison Kentucky
BEREA
Carduus nutans
BEREA002864 Madison Kentucky
BEREA
Carduus nutans
31973000005456 Gallatin Kentucky 09 Jun 1984 Thieret, John
55063
KNK
Carduus nutans
TENN-V-0237403 Sequatchie Tennessee
TENN
Carduus nutans
TENN-V-0237407 Shelby Tennessee
TENN
Carduus nutans
TENN-V-0237400 Rutherford Tennessee
TENN
Carduus nutans
TENN-V-0237376 Davidson Tennessee
TENN
Carduus nutans
TENN-V-0237395 Van Buren Tennessee
TENN
Carduus nutans
TENN-V-0241521 Hamilton Tennessee
TENN
Carduus nutans
TENN-V-0237445 Anderson Kentucky
TENN
Carduus nutans
31234100244720 Clinton Kentucky 25 Jul 1999 Mark A. Gorton
99-806
EKY
Carduus nutans
31234100244894 Muhlenberg Kentucky 02 Jun 1993 Landon E. McKinney
5643
EKY
Carduus nutans
31234100244860 Marion Kentucky 18 Jun 1994 M. H. McGuffey
49
EKY
Carduus nutans
31234100244852 Madison Kentucky 23 Jun 1978 J. Stuart Lassetter
2965
EKY
Carduus nutans
31234100244837 Madison Kentucky 30 Jun 1993 Timothy J. Weckman
401
EKY
Carduus nutans
31234100244753 Franklin Kentucky 11 Jun 1957 Mary E. Wharton
10842
EKY
Carduus nutans
31234100244845 Madison Kentucky 20 Jun 1975 J. Stuart Lassetter
2420
EKY
Carduus nutans
31234100244787 Jefferson Kentucky 22 Jun 2005 Patricia D. Haragan
25167
EKY
Carduus nutans
31234100244779 Hickman Kentucky 12 Jul 1984 Raymond Athey
4938
EKY
Carduus nutans
BEREA002863 Madison Kentucky
BEREA
Carduus nutans
TENN-V-0237381 Giles Tennessee
TENN
Carduus nutans
TENN-V-0237396 Williamson Tennessee
TENN
Carduus nutans
TENN-V-0237380 Franklin Tennessee
TENN
Carduus nutans
TENN-V-0237384 Henderson Tennessee
TENN
Carduus nutans
TENN-V-0237378 Dyer Tennessee
TENN
Carduus nutans
31234100244696 Anderson Kentucky 14 Jun 1978 William S. Bryant
461
EKY
Carduus nutans
31234100244746 Fayette Kentucky Danny Husband
190
EKY
Carduus nutans
31234100244829 Madison Kentucky 12 Jun 1993 D. Collinsworth
45
EKY
Carduus nutans
31234100244795 Logan Kentucky 12 Jun 1976 Raymond Athey
3415
EKY
Carduus nutans
BEREA002867 Cannon Tennessee
BEREA
Carduus nutans
BEREA002858 Garrard Kentucky
BEREA
Carduus nutans
31973000005454 Campbell Kentucky 03 Jun 1982 Buddell, George
616
KNK
Carduus nutans
TENN-V-0237383 Hamblen Tennessee
TENN
Carduus nutans
TENN-V-0237410 White Tennessee
TENN
Carduus nutans
TENN-V-0237440 Clark Kentucky
TENN
Carduus nutans
TENN-V-0237446 Warren Kentucky
TENN
Carduus nutans
TENN-V-0237390 Loudon Tennessee
TENN
Carduus nutans
TENN-V-0237402 Rutherford Tennessee
TENN
Carduus nutans
APSC0121905 Oldham Kentucky
APSC
Carduus nutans
APSC0074191 Warren Kentucky
APSC
Carduus nutans
31234100757390 Barren Kentucky 09 Jun 1979 George P. Johnson
741
EKY
Carduus nutans
31234100244712 Clark Kentucky 20 Jun 1978 Elizabeth M. Browne
78D1.1
EKY
Carduus nutans
BEREA002868 DeKalb Tennessee
BEREA
Carduus nutans
BEREA002860 Madison Kentucky
BEREA
Carduus nutans
BEREA002861 Madison Kentucky
BEREA
Carduus nutans
31973000005457 Kenton Kentucky 24 Jun 1978 Applegarth, Patrick
357
KNK
Carduus nutans
MDKY00010313 Clark Kentucky
MDKY
Carduus nutans
MDKY00010316 Fleming Kentucky
MDKY
Carduus nutans
TENN-V-0237398 Rutherford Tennessee
TENN
Carduus nutans
TENN-V-0237379 Grundy Tennessee
TENN
Carduus nutans
TENN-V-0237409 Unicoi Tennessee
TENN
Carduus nutans
TENN-V-0237404 Sequatchie Tennessee
TENN
Carduus nutans
TENN-V-0237391 Macon Tennessee
TENN
Carduus nutans
TENN-V-0237399 Rutherford Tennessee
TENN
Carduus nutans
TENN-V-0237408 Sullivan Tennessee
TENN
Carduus nutans
APSC0027226 Stewart Tennessee
APSC
Carduus nutans
31234100244886 Montgomery Kentucky 28 Jun 1984 Julian Campbell
521
EKY
Carduus nutans
31234100244878 Mercer Kentucky 28 Jun 1976 Donald Pelly
3
EKY
Carduus nutans
BEREA002866 Rockcastle Kentucky
BEREA
Carduus nutans
31973000005455 Carroll Kentucky 09 Jun 1984 Thieret, John
55064
KNK
Carduus nutans
MDKY00010315 Rowan Kentucky
MDKY
Carduus nutans
TENN-V-0237441 Logan Kentucky
TENN
Carduus nutans
TENN-V-0237406 Shelby Tennessee
TENN
Carduus nutans
TENN-V-0237374 Carroll Tennessee
TENN
Carduus nutans
UCHT007920 Benton Tennessee 17 Jun 1998 Don Sudbrink
UCHT
Carduus nutans
APSC0108275 Nelson Kentucky
APSC
Carduus nutans
APSC0027234 Montgomery Tennessee
APSC
Carduus nutans
APSC0074160 Warren Kentucky
APSC
Carduus nutans
APSC0027227 Montgomery Tennessee
APSC
Carduus nutans
APSC0085851 Jefferson Kentucky
APSC
Carduus nutans
APSC0074159 Warren Kentucky
APSC
Carduus nutans
APSC0027233 Stewart Tennessee
APSC
Carduus nutans
31234100244803 Madison Kentucky 16 Jun 1993 Gary W. Libby
422
EKY
Carduus nutans
ETSU007991 Knox Tennessee
ETSU
Carduus nutans
TENN-V-0237386 Jackson Tennessee
TENN
Carduus nutans
TENN-V-0237392 Maury Tennessee
TENN
Carduus nutans
TENN-V-0237382 Greene Tennessee
TENN
Carduus nutans
TENN-V-0237397 Wilson Tennessee
TENN
Carduus nutans
TENN-V-0237375 Davidson Tennessee
TENN
Carduus nutans
TENN-V-0237393 Maury Tennessee
TENN
Carduus nutans
APSC0050758 Larue Kentucky 26 May 1993 Landon McKinney
APSC
Carduus nutans
APSC0027220 Trigg Kentucky
APSC
Carduus nutans
HTTU002700 Hamilton Tennessee
HTTU
Carduus nutans
HTTU002695 Fentress Tennessee
HTTU
Carduus nutans
HTTU002690 Tennessee
HTTU
Carduus nutans
HTTU002689 Putnam Tennessee
HTTU
Carduus nutans
HTTU002693 Williamson Tennessee
HTTU
Carduus nutans
HTTU028911 White Tennessee
HTTU
Carduus nutans
MTSU021340 Wilson Tennessee 01 May 1987 Fred Funte
4
MTSU
Carduus nutans
MTSU021328 Rutherford Tennessee 25 Apr 1985 Kenneth Hayes
MTSU
Carduus nutans
MTSU021325 Rutherford Tennessee 05 Jun 1984 R. J. Beel
7
MTSU
Carduus nutans
MTSU021324 Rutherford Tennessee 16 Jun 1981 Jeff Peery
13
MTSU
Carduus nutans
MTSU021322 Rutherford Tennessee 06 Jul 1981 Joao M. C. Ribeiro
41
MTSU
Carduus nutans
MTSU021341 Rutherford Tennessee 20 Apr 1992 Theresa DePalo
22
MTSU
Carduus nutans
MTSU021348 Davidson Tennessee 22 Apr 1997 Josh Rogers
32
MTSU
Carduus nutans
MTSU021332 Davidson Tennessee 29 Apr 1986 Bill Mastalerz
33
MTSU
Carduus nutans
MTSU021337 Rutherford Tennessee 29 Apr 1990 Hope Dodson
46
MTSU
Carduus nutans
MTSU021338 Rutherford Tennessee 21 Apr 1991 Gina Rigney
26
MTSU
Carduus nutans
MTSU021349 Rutherford Tennessee 29 Apr 1999 Lori Bruner
48
MTSU
Carduus nutans
MTSU021347 Knox Tennessee 20 Apr 1997 Lora Carlson
42
MTSU
Carduus nutans
MTSU021352 Giles Tennessee 01 May 2004 Lindsey Jett
48
MTSU
Carduus nutans
MTSU021318 Rutherford Tennessee 24 Jun 1994 Terri Hogan
178-94
MTSU
Carduus nutans
31234100244761 Franklin Kentucky 09 Jul 1956 Mary E. Wharton
10256
EKY
Carduus nutans
BEREA002859 Garrard Kentucky
BEREA
Carduus nutans
TENN-V-0237394 Putnam Tennessee
TENN
Carduus nutans
TENN-V-0237388 Knox Tennessee
TENN
Carduus nutans
TENN-V-0237387 Jefferson Tennessee
TENN
Carduus nutans
TENN-V-0237371 Benton Tennessee 17 Jun 1998 Don Sudbrink
TENN
Carduus nutans
TENN-V-0237373 Campbell Tennessee
TENN
Carduus nutans
TENN-V-0237405 Sevier Tennessee
TENN
Carduus nutans
000857 Carroll Tennessee 01 Jun 1970 Betty Ann Thompson
13
BOON
Carduus nutans
APSC0069899 Shelby Kentucky
APSC
Carduus nutans
APSC0027237 Montgomery Tennessee
APSC
Carduus nutans
APSC0027235 Houston Tennessee
APSC
Carduus nutans
UCHT029716 Hamilton Tennessee
UCHT
Carduus nutans
UCHT029714 DeKalb Tennessee
UCHT
Carduus nutans
UCHT029715 Marion Tennessee
UCHT
Carduus nutans
UCHT039724 Grundy Tennessee
UCHT
Carduus nutans
MTSU021319 Davidson Tennessee 09 May 2001 Angela Cathey
165
MTSU
Carduus nutans
MTSU021342 Rutherford Tennessee 27 Apr 1992 M. Derek Goodman
46
MTSU
Carduus nutans
MTSU021334 Coffee Tennessee 27 Apr 1987 Kim A. Davis
35
MTSU
Carduus nutans
MTSU021354 Maury Tennessee 26 Apr 2007 Jill Price
45
MTSU
Carduus nutans
MTSU021330 Rutherford Tennessee 12 Apr 1986 Jeri Johnson
MTSU
Carduus nutans
MTSU021336 Davidson Tennessee 27 Apr 1986 Monteia Moore
39
MTSU
Carduus nutans
MTSU021346 Cannon Tennessee 25 Apr 1997 Ashley Drake
21
MTSU
Carduus nutans
MTSU021344 Rutherford Tennessee 29 Apr 1995 Kenya Beverly
44
MTSU
Carduus nutans
MTSU021351 Rutherford Tennessee 24 Apr 2000 Mary Etta Boulden
50
MTSU
Carduus nutans
MTSU021321 Davidson Tennessee 20 Jun 1981 James Bonner
12
MTSU
Carduus nutans
MTSU021350 Williamson Tennessee 30 Apr 2000 Melanie Messina
20
MTSU
Carduus nutans
MTSU021345 Rutherford Tennessee 01 May 1995 Cristina Scola
32
MTSU
Carduus nutans
TENN-V-0243817 Knox Tennessee
TENN
Carduus nutans
TENN-V-0237385 Henderson Tennessee
TENN
Carduus nutans
UOS00715 Franklin Tennessee 05 Jun 2007 Mary P. Priestley
UOS
Carduus nutans
UOS00713 Franklin Tennessee 06 Jun 1996 Douglas E. Kennemore, Jr.
UOS
Carduus nutans
UOS00714 Franklin Tennessee 06 Jun 1996 Douglas E. Kennemore, Jr.
UOS
Carduus nutans
31234100244704 Barren Kentucky 27 May 1993 Charles J. Lapham
48
EKY
Carduus nutans
31234100389640 Madison Kentucky 11 Oct 1993 Gary W. Libby
190
EKY
Carduus nutans
BEREA022919 Monroe Kentucky
BEREA
Carduus nutans
BEREA002857 Clark Kentucky
BEREA
Carduus nutans
MDKY00010314 Logan Kentucky
MDKY
Carduus nutans
TENN-V-0237377 DeKalb Tennessee
TENN
Carduus nutans
TENN-V-0237389 Loudon Tennessee
TENN
Carduus nutans
TENN-V-0237401 Rutherford Tennessee
TENN
Carduus nutans
APSC0121906 Nelson Kentucky
APSC
Carduus nutans
APSC0027229 Smith Tennessee
APSC
Carduus nutans
APSC0027228 Stewart Tennessee
APSC
Carduus nutans
APSC0027219 Stewart Tennessee
APSC
Carduus nutans
APSC0074194 Barren Kentucky
APSC
Carduus nutans
APSC0074156 Warren Kentucky
APSC
Carduus nutans
APSC0027223 Davidson Tennessee
APSC
Carduus nutans
APSC0074158 Warren Kentucky
APSC
Carduus nutans
APSC0027236 Montgomery Tennessee
APSC
Carduus nutans
APSC0027218 Stewart Tennessee
APSC
Carduus nutans
APSC0107389 Trimble Kentucky
APSC
Carduus nutans
APSC0074157 Warren Kentucky
APSC
Carduus nutans
SWMT04196 Williamson Tennessee 12 Jun 1982 Rick Sullivan
SWMT
Carduus nutans
BEREA025750 Madison Kentucky
BEREA
Carduus nutans
BEREA025751 Madison Kentucky
BEREA
Carduus nutans
TENN-V-0237372 Bledsoe Tennessee
TENN
Carduus nutans
APSC0027224 Montgomery Tennessee
APSC
Carduus nutans
APSC0027232 Franklin Tennessee
APSC
Carduus nutans
APSC0080195 Gallatin Kentucky
APSC
Carduus nutans
APSC0074192 Marshall Kentucky
APSC
Carduus nutans
APSC0027225 Christian Kentucky 10 Jun 1968
APSC
Carduus nutans
ETSU011763 Washington Tennessee
ETSU
Carduus nutans
ETSU011766 Sullivan Tennessee
ETSU
Carduus nutans
ETSU011764 Washington Tennessee
ETSU
Carduus nutans
ETSU011762 Washington Tennessee
ETSU
Carduus nutans
ETSU011765 Washington Tennessee
ETSU
Carduus nutans
HTTU002698 Maury Tennessee
HTTU
Carduus nutans
HTTU002691 Williamson Tennessee
HTTU
Carduus nutans
HTTU002699 Haywood Tennessee
HTTU
Carduus nutans
HTTU002697 Wilson Tennessee
HTTU
Carduus nutans
HTTU002692 Williamson Tennessee
HTTU
Carduus nutans
HTTU002688 DeKalb Tennessee
HTTU
Carduus nutans
MEM007077 Marshall Kentucky 05 Jun 1972 Raymond Athey
1794
MEM
Carduus nutans
MEM003427 Scott Kentucky 08 Jun 1961 William Wayman
s.n.
MEM
Carduus nutans
MEM009451 Warren Kentucky 08 Jun 1971 Kenneth A. Nicely
3256
MEM
Carduus nutans
TENN-V-0243810 Knox Tennessee 07 Jun 2016 B.J. Crain
91
TENN
Carduus nutans
HTTU002701 Sumner Tennessee
HTTU
Carduus nutans
HTTU002685 Smith Tennessee
HTTU
Carduus nutans
HTTU002694 Tennessee
HTTU
Carduus nutans
HTTU002686 Sumner Tennessee
HTTU
Carduus nutans
HTTU002684 Putnam Tennessee
HTTU
Carduus nutans
HTTU002687 DeKalb Tennessee
HTTU
Carduus nutans
HTTU002683 DeKalb Tennessee
HTTU
Carduus nutans
MTSU021353 Franklin Tennessee 19 Apr 2005 Karen Williams
60
MTSU
Carduus nutans
MTSU021335 Davidson Tennessee 23 Apr 1991 Brian White
28
MTSU
Carduus nutans
MTSU021343 Rutherford Tennessee 15 Apr 1992 Mary Penuel
39
MTSU
Carduus nutans
MTSU021329 Davidson Tennessee 19 Apr 1986 Jim Ohlsson
11
MTSU
Carduus nutans
MTSU021323 Rutherford Tennessee 15 Jun 1981 Teri Coats
11
MTSU
Carduus nutans
MTSU021339 Rutherford Tennessee 27 Apr 1986 Benedicta Kurunwune
47
MTSU
Carduus nutans
MTSU021331 Rutherford Tennessee 08 Apr 1986 Jeri Johnson
11
MTSU
Carduus nutans
MTSU021333 Rutherford Tennessee
MTSU
Carduus nutans
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Plant Photos

Carduus nutans - Ashley B. Morris
Carduus nutans - Ashley B. Morris
Carduus nutans - Dwayne Estes
Carduus nutans - Dwayne Estes
Carduus nutans - Dwayne Estes
Carduus nutans - Dwayne Estes
Carduus nutans - Joey Shaw
Carduus nutans - Joey Shaw
Carduus nutans - Joey Shaw
Carduus nutans - Joey Shaw
Carduus nutans - Joey Shaw
Carduus nutans - Joey Shaw
Carduus nutans - Theo Witsell
Carduus nutans - Theo Witsell
Carduus nutans - Theo Witsell
Carduus nutans - Theo Witsell
Carduus nutans - Theo Witsell
Carduus nutans - Theo Witsell