Sceptridium biternatum

Classification	Pteridophytes
Family	Ophioglossaceae
Family (Traditional)	Ophioglossaceae
Genus	Sceptridium
Species:	Sceptridium biternatum (Savigny) Lyon
Common Name	Sparse Lobed Grape Fern
Growth Habit	fern
Plant Notes	**
Characteristics	Armaments: No; Milky Sap: No; Photosynthetic: Yes; Stipules: No; Woody Herbaceous: Herbaceous; Natural Area Presence: DRU; Natural Area Presence: GILES; Natural Area Presence: GM; Natural Area Presence: GRSM; Natural Area Presence: LC; Natural Area Presence: LGS; Natural Area Presence: PK; Natural Area Presence: RCSP; Natural Area Presence: UCR; Duration: Perennial; General Ecology: Terrestrial; Growth Habit: fern; Moisture Requirements: Dry-mesic; Moisture Requirements: Mesic; Moisture Requirements: Mesic-Hydric; Moisture Requirements: Hydric; Moisture Requirements: Wetland; Region of Origin: North America; Reproductive Phenology: August; Reproductive Phenology: September; Reproductive Phenology: October; Leaf Arrangement: basal; Leaf Complexity: compound; Leaf Dissection: 2; Leaf Dissection: 2.5; Leaf Dissection: 3; Leaf Fragrance: No; Leaf Persistance: Deciduous; Leaf Persistance: Tardily Deciduous; Leaf Petiole Type: petiolate; Leaf Rank: N/A; Leaf Type: fern-like; Leaf Venation: pinnate; Leaflet Margins: serrate; Leaflet Number: 3; Leaflet Number: 4; Leaflet Number: 5; Leaflet Number: 6; Leaflet Number: 7; Leaflet Number: 8; Leaflet Number: 9; Anther Type: N/A; Carpel Number/Pistil: N/A; Fruit Complexity: N/A; Fruit Type: N/A; Fruit Wet/Dry: N/A; Hypanthium Present: No; Inflorescence Position: N/A; Inflorescence Types: N/A; Ovary Position: N/A; Overall Flower Symmetry: N/A; Ovules per Carpel: N/A; Perianth Present: No; Petal Fusion: N/A; Petal Number: N/A; Pistil Complexity: N/A; Pistil Number: N/A; Placentation Type: N/A; Sepal Fusion: N/A; Sepal Number: N/A; Sexuality: N/A; Sporangium Arrangement: underground sporangia; Stamen Number: N/A; Tepal Number: N/A
C value	5
Status	Native; AGCP (FAC), EMP (FAC) (NWPL)
Specimen(s)	View and Download Specimen Details »

** Not applicable or data not available.

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Species Distribution Map

This species has been reported in the counties highlighted in the interactive distribution map below. Click on a county to display the its name. Click the question mark icon () to view additional information about the ecoregions shown. In the map legend, there are links to toggle the visibility of the ecoregions and/or their labels. In addition, clicking on an ecoregion name will highlight that region on the map.

State Boundaries
Counties
Counties - Species Present

Ecoregions

Southeastern Plains
- Blackland Prarie
Blue Ridge Mountains
Ridge and Valley
Cumberland Plateau
Cumberland Mountains
Allegheny Plateau

Interior Plateau
- Bluegrass Region
- Nashville Basin
- Pennyroyal Karst Plain
- Western Highland Rim
- Eastern Highland Rim

Interior River Valleys and Hills
- Shawnee / Caseyville Hills
Mississippi Alluvial Plain
Mississippi Valley Loess Plains
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Synonyms

Synonym	Full Citation
Botrychium biternatum	Botrychium biternatum (Sav.) Underw.
Botrychium obliquum var. oblongifolium	Botrychium obliquum var. oblongifolium E. W. Graves
Botrychium tenuifolium	Botrychium tenuifolium

Specimens and Distribution

This species has been reported in the following counties by the herbaria listed. An overview of the individual specimens are provided in the table that follows. Click on the accession number to view details; click on column headers to sort.

Counties included on distribution map: Boyle (KY), Carter (KY), Elliott (KY), Hancock (KY), Laurel (KY), Lyon (KY), Madison (KY), Menifee (KY), Powell (KY), Robertson (KY), Rowan (KY), Wolfe (KY), Blount (TN), Bradley (TN), Campbell (TN), Carter (TN), Cheatham (TN), Chester (TN), Cocke (TN), Crockett (TN), Cumberland (TN), Davidson (TN), Decatur (TN), Dickson (TN), Fayette (TN), Franklin (TN), Gibson (TN), Giles (TN), Hamilton (TN), Hancock (TN), Hardeman (TN), Henderson (TN), Henry (TN), Hickman (TN), Humphreys (TN), Jackson (TN), Johnson (TN), Knox (TN), Lewis (TN), McMinn (TN), Macon (TN), Marion (TN), Maury (TN), Monroe (TN), Montgomery (TN), Morgan (TN), Obion (TN), Overton (TN), Perry (TN), Polk (TN), Putnam (TN), Rhea (TN), Roane (TN), Rutherford (TN), Sequatchie (TN), Sevier (TN), Shelby (TN), Sullivan (TN), Sumner (TN), Unicoi (TN), Union (TN), Warren (TN), Washington (TN), Wayne (TN), Weakley (TN), White (TN), Wilson (TN)

Herbaria represented by specimen data listed below: MDKY, UOS, ETSU, EKY, HTTU, KNK, UCHT, MEM, SWMT, VDB, BEREA, TENN

Range of years during which specimens were collected: 1935 - 2019

Accession No. (click for full details)	County	State	Coll. Date	Collector & Collection No.	Herbarium & Herbarium Name Used
etsu005212	Unicoi	Tennessee	16 Sep 2010	Frosty Levy	ETSU Botrychium biternatum
MDKY00000131	Rowan	Kentucky			MDKY Botrychium biternatum
MDKY00000129	Wolfe	Kentucky			MDKY Botrychium biternatum
MDKY00000127	Rowan	Kentucky			MDKY Botrychium biternatum
TENN-V-0002033	Cumberland	Tennessee	26 Oct 1962	James S. Pringle NO. 31259	TENN Botrychium biternatum
TENN-V-0002048	Hickman	Tennessee	13 May 1998	Dwayne Estes 183	TENN Botrychium biternatum
TENN-V-0002121	Warren	Tennessee	02 Oct 1943	Jesse M. Shaver 4367	TENN Botrychium biternatum
TENN-V-0171086	Gibson	Tennessee			TENN Botrychium biternatum
TENN-V-0002043	Hamilton	Tennessee	05 Sep 1952	Jesse M. Shaver 10928	TENN Botrychium biternatum
TENN-V-0002107	Sequatchie	Tennessee	05 Sep 1945	Jessee M. Shaver 8248	TENN Botrychium biternatum
TENN-V-0002094	Montgomery	Tennessee	22 Sep 1949	Jesse M. Shaver 10549	TENN Botrychium biternatum
TENN-V-0002117	Unicoi	Tennessee	24 Aug 1941	Jesse M. Shaver 7896B	TENN Botrychium biternatum
TENN-V-0002041	Giles	Tennessee	19 Jul 2003	Dwayne Estes 5030	TENN Botrychium biternatum
TENN-V-0002108	Sequatchie	Tennessee	26 Dec 2006	John Evans 49	TENN Botrychium biternatum
31234100021060	Powell	Kentucky	06 Aug 2001	Timothy J. Weckman 6688	EKY Botrychium biternatum
MDKY00000137	Rowan	Kentucky			MDKY Botrychium biternatum
MDKY00000128	Rowan	Kentucky			MDKY Botrychium biternatum
MDKY00000132	Rowan	Kentucky			MDKY Botrychium biternatum
TENN-V-0002111	Sullivan	Tennessee	24 Sep 1938	Herman O'Dell s.n	TENN Botrychium biternatum
TENN-V-0002122	Warren	Tennessee	04 May 1947	Jesse M. Shaver 9728	TENN Botrychium biternatum
TENN-V-0002029	Crockett	Tennessee	17 Sep 1947	Jesse M Shaver 9996	TENN Botrychium biternatum
TENN-V-0002609	Weakley	Tennessee			TENN Botrychium biternatum
TENN-V-0002036	Fayette	Tennessee	19 Oct 1972	H. R. DoSeln, Mr. Pope	TENN Botrychium biternatum
TENN-V-0002104	Roane	Tennessee	26 Aug 1974	S.F. Hale, F.E. Williams 48617	TENN Botrychium biternatum
TENN-V-0002098	Overton	Tennessee	01 Sep 1943	Jamison and Shaver 4102	TENN Botrychium biternatum
TENN-V-0002046	Henderson	Tennessee	16 Sep 1952	Jesse M. Shaver 10887	TENN Botrychium biternatum
TENN-V-0002096	Morgan	Tennessee	05 Oct 1967	A. M. Evans 3404	TENN Botrychium biternatum
TENN-V-0002031	Cumberland	Tennessee	14 Nov 1978	A M Evans, B E Wofford	TENN Botrychium biternatum
N/A	Elliott	Kentucky	15 Mar 2012	M. Greene	EKY Botrychium biternatum
BEREA000437	Laurel	Kentucky			BEREA Sceptridium biternatum
MDKY00000136	Rowan	Kentucky			MDKY Botrychium biternatum
TENN-V-0002042	Hamilton	Tennessee	00 June 1971	Frank Bowers	TENN Botrychium biternatum
TENN-V-0002044	Hardeman	Tennessee	21 Apr 1946	Jesse M. Shaver 9169	TENN Botrychium biternatum
TENN-V-0002100	Polk	Tennessee	30 Mar 1956	A. J. Sharp 20406	TENN Botrychium biternatum
TENN-V-0002110	Shelby	Tennessee	16 Sep 1952	Jesse M. Shaver 10888	TENN Botrychium biternatum
TENN-V-0002090	Montgomery	Tennessee	18 Nov 1945	Jesse M. Shaver 8148	TENN Botrychium biternatum
TENN-V-0002113	Sumner	Tennessee	03 Nov 1946	Jesse M. Shaver 9820	TENN Botrychium biternatum
TENN-V-0002032	Cumberland	Tennessee	16 Aug 1951	FHN, AJS 16205	TENN Botrychium biternatum
TENN-V-0002040	Giles	Tennessee	23 Aug 2003	Dwayne Estes 5389	TENN Botrychium biternatum
UCHT000278	Hamilton	Tennessee	30 Aug 2002	T. Govus, R. Evans 3961	UCHT Sceptridium biternatum
UCHT000273		Tennessee	01 Oct 1981	Greg Carlson	UCHT Sceptridium biternatum
UCHT000271	Marion	Tennessee	07 Aug 1998	John Beck 2707	UCHT Sceptridium biternatum
31234100739679	Hancock	Kentucky	17 Sep 1995	Timothy J. Weckman 2039	EKY Botrychium biternatum
31234100779857	Menifee	Kentucky	26 Jul 2017	Thomas McFadden 600	EKY Sceptridium biternatum
TENN-V-0002128	Henry	Tennessee	13 Sep 1952	Jesse M. Shaver 10898	TENN Botrychium biternatum
TENN-V-0002039	Giles	Tennessee	23 Aug 2003	Dwayne Estes 5436	TENN Botrychium biternatum
TENN-V-0002017	Sumner	Tennessee	03 Nov 1946	Jesse M. Shaver 9820 B	TENN Botrychium biternatum
TENN-V-0002027	Chester	Tennessee	17 Sep 1952	Jesse M. Shaver 10882	TENN Botrychium biternatum
TENN-V-0002112	Johnson	Tennessee	08 Sep 1946	Jesse M. Shaver 9747	TENN Botrychium biternatum
TENN-V-0002025	Blount	Tennessee	07 Aug 1971	Jerry C. Odenwelder and C.A. Boyll 71-181	TENN Botrychium biternatum
TENN-V-0002103	Roane	Tennessee	21 Aug 1981	Thomas S. Patrick 3581	TENN Botrychium biternatum
TENN-V-0002116	Unicoi	Tennessee	24 Aug 1941	Jesse M. Shaver 7896A	TENN Botrychium biternatum
TENN-V-0002056	Monroe	Tennessee	23 Jul 1977	Jeffry L. Malter 53598	TENN Botrychium biternatum
UCHT000274	Hamilton	Tennessee	08 Sep 1991	Gary H. Marton 8987	UCHT Sceptridium biternatum
BRIT188212	Humphreys	Tennessee	25 Sep 1991	R. Kral 79846	VDB Botrychium biternatum
TENN-V-0002106	Rutherford	Tennessee	14 Oct 2001	Dwayne Estes 3009	TENN Botrychium biternatum
TENN-V-0002126	Wayne	Tennessee	12 Sep 1945	Jesse M. Shaver 8283	TENN Botrychium biternatum
TENN-V-0002131	White	Tennessee	05 Sep 1937	Jesse M. Shaver 2180	TENN Botrychium biternatum
TENN-V-0002034	Decatur	Tennessee	13 Oct 1949	EHC, FWW, RES 14827	TENN Botrychium biternatum
TENN-V-0002130	White	Tennessee	21 Sep 1979	Thomas S. Patrick 2629	TENN Botrychium biternatum
TENN-V-0002097	Morgan	Tennessee	07 Oct 1966	A.M. Evans 2259	TENN Botrychium biternatum
TENN-V-0002101	Rhea	Tennessee	13 Sep 1937	Jesse M. Shaver 2177	TENN Botrychium biternatum
UCHT000258	Marion	Tennessee	16 Oct 2002	erin king 387	UCHT Sceptridium biternatum
UCHT000259	Hamilton	Tennessee	01 Oct 1981	Angela May S.N.	UCHT Sceptridium biternatum
HTTU028583	Putnam	Tennessee			HTTU Sceptridium biternatum
BRIT188219	Perry	Tennessee	24 Apr 1972	R. Kral 45673	VDB Botrychium biternatum
BRIT188217	Maury	Tennessee	12 May 1946	Jesse M. Shaver 9213	VDB Botrychium biternatum
BRIT188206	Davidson	Tennessee	05 Oct 1952	E. Quarterman 4364	VDB Botrychium biternatum
TENN-V-0002127	Weakley	Tennessee	13 Sep 1952	Jesse M. Shaver 10899A	TENN Botrychium biternatum
TENN-V-0002118	Unicoi	Tennessee	10 Sep 1936	Jesse M. Shaver 2188	TENN Botrychium biternatum
TENN-V-0002091	Obion	Tennessee	23 Aug 1971	A.M. Evans, Gene Wofford	TENN Botrychium biternatum
TENN-V-0002125	Washington	Tennessee	08 Sep 1946	Jesse M. Shaver 9777	TENN Botrychium biternatum
TENN-V-0002123	Washington	Tennessee	30 Jul 1950	RES & AJS 7467	TENN Botrychium biternatum
UCHT000265		Tennessee	14 Sep 1982	D. Dubase, B. Cheever	UCHT Sceptridium biternatum
UCHT000264	Hamilton	Tennessee	00 June 1971	Frank Bowers	UCHT Sceptridium biternatum
HTTU000325	Jackson	Tennessee			HTTU Botrychium biternatum
TENN-V-0244976	Knox	Tennessee	19 Nov 2019	Jessica M. Budke 301	TENN Sceptridium biternatum
UOS08645	Franklin	Tennessee	02 Oct 1997	Mary P. Priestley	UOS Sceptridium biternatum
BRIT188215	Marion	Tennessee	30 Aug 1936	Jesse M. Shaver 2192	VDB Botrychium biternatum
BRIT188204	Cumberland	Tennessee	31 Aug 1989	V.E. McNeilus 89-881	VDB Botrychium biternatum
BRIT188214	Macon	Tennessee	23 Sep 1981	R. Kral 67898	VDB Botrychium biternatum
31234100783354	Madison	Kentucky	15 Sep 2016	Thomas McFadden 271	EKY Sceptridium biternatum
MDKY00000130	Rowan	Kentucky			MDKY Botrychium biternatum
MDKY00000135	Carter	Kentucky			MDKY Botrychium biternatum
MDKY00000133	Rowan	Kentucky			MDKY Botrychium biternatum
TENN-V-0002026	Campbell	Tennessee	20 Sep 2012	Aaron Floden & Peter Zale 2267	TENN Botrychium biternatum
TENN-V-0002037	Franklin	Tennessee	16 Sep 1937	2161	TENN Botrychium biternatum
TENN-V-0002129	White	Tennessee	05 Sep 1937	Jesse M. Shaver 2182	TENN Botrychium biternatum
TENN-V-0002092	Montgomery	Tennessee	29 Sep 1979	Edward W. Chester 4312	TENN Botrychium biternatum
TENN-V-0002095	Montgomery	Tennessee	27 Feb 1944	R. E. Shanks 2678	TENN Botrychium biternatum
TENN-V-0002102	Roane	Tennessee	08 Sep 1948	Jesse M. Shaver 10392C	TENN Botrychium biternatum
TENN-V-0002115	Unicoi	Tennessee	24 Aug 1941	Jesse M. Shaver 7896C	TENN Botrychium biternatum
UOS08642	Franklin	Tennessee	01 Jun 1952	Jean A. Yeatman	UOS Sceptridium biternatum
SWMT05481	Shelby	Tennessee	12 Oct 2002	Michael Burke 909	SWMT Botrychium biternatum
SWMT00060		Tennessee	14 May 1937	Boyles	SWMT Botrychium tenuifolium
TENN-V-0002045	Hardeman	Tennessee	14 Sep 1949	R.E. Shanks, E.H. Cooley, & F.W. Woods 9241	TENN Botrychium biternatum
TENN-V-0002049	Knox	Tennessee	26 Nov 1978	Mrs. Curtis Shelton	TENN Botrychium biternatum
UCHT000275	Hamilton	Tennessee	01 Oct 1981	Mark Weehard 12	UCHT Sceptridium biternatum
UCHT000268	Hamilton	Tennessee	14 Sep 1982	McKee, Hoosier s.n	UCHT Sceptridium biternatum
UCHT000276	Hamilton	Tennessee	13 Nov 1985	Herman Rada	UCHT Sceptridium biternatum
BRIT188213	Lewis	Tennessee	10 Aug 1945	C. King 390	VDB Botrychium biternatum
BRIT188222	Wayne	Tennessee	10 Oct 1978	R. Kral 62863	VDB Botrychium biternatum
BRIT188218	Monroe	Tennessee	08 Sep 1952	Jesse M. Shaver 10907	VDB Botrychium dissectum
BRIT188203	Cheatham	Tennessee	15 Oct 1979	R. Kral 64696	VDB Botrychium biternatum
MEM007960	Shelby	Tennessee	04 Sep 1970	Elizabeth M. Browne, Edward T. Browne Jr. 70M2.5	MEM Botrychium biternatum
UCHT000260	Bradley	Tennessee	03 Jun 1989	D. Houck 2665	UCHT Sceptridium biternatum
UCHT000272	Hamilton	Tennessee	03 Sep 1981	Myron Donesky	UCHT Sceptridium biternatum
UCHT000262	Marion	Tennessee	22 Mar 1998	John Beck 1355	UCHT Sceptridium biternatum
31973000004402	Boyle	Kentucky	03 Aug 1979	Thieret, John 51133	KNK Botrychium biternatum
TENN-V-0002119	Unicoi	Tennessee	07 Sep 1949	R.E. Shanks and F.W. Woods and E.H. Cooley 8864	TENN Botrychium biternatum
TENN-V-0002038	Gibson	Tennessee	13 Sep 1952	Jesse M. Shaver 10901	TENN Botrychium biternatum
TENN-V-0002035	Dickson	Tennessee	30 Sep 1945	Jesse M. Shaver 8237	TENN Botrychium biternatum
TENN-V-0002028	Crockett	Tennessee	17 Sep 1947	Jesse M. Shaver 9998	TENN Botrychium biternatum
TENN-V-0002124	Washington	Tennessee	05 Sep 1952	Jesse M. Shaver 10925	TENN Botrychium biternatum
TENN-V-0002089	Overton	Tennessee	13 Sep 1994	V. McNeilus 94-990	TENN Botrychium biternatum
TENN-V-0002030	Crockett	Tennessee	17 Sep 1947	Jesse M. Shaver 9998B	TENN Botrychium biternatum
TENN-V-0244884	Hamilton	Tennessee	20 Oct 2019	A. Workinger s.n.	TENN Sceptridium biternatum
BRIT188216	Marion	Tennessee	22 Oct 1966	D.P. Whittier 101472	VDB Botrychium biternatum
ETSU032237	Carter	Tennessee		James T Donaldson	ETSU Sceptridium biternatum
ETSU031749	Washington	Tennessee			ETSU Sceptridium biternatum
TENN-V-0002109	Shelby	Tennessee	16 Sep 1952	Jesse M. Shaver 10888	TENN Botrychium biternatum
TENN-V-0002114	Sumner	Tennessee	07 Oct 1949	RES, FWW, EHC 14268	TENN Botrychium biternatum
MDKY00000134	Rowan	Kentucky			MDKY Botrychium biternatum
31234100021169	Robertson	Kentucky	15 Aug 2002	Timothy J. Weckman 7668	EKY Botrychium biternatum
TENN-V-0002120	Union	Tennessee	04 Sep 1952	Jesse M. Shaver 10934	TENN Botrychium biternatum
TENN-V-0002105	Roane	Tennessee	08 Sep 1948	Jesse M. Shaver 10392B	TENN Botrychium biternatum
BRIT188202	Campbell	Tennessee	04 Sep 1952	Jesse M. Shaver 10931	VDB Botrychium biternatum
BRIT188211	Hardeman	Tennessee	01 Jun 1952	Jesse M. Shaver 10955	VDB Botrychium dissectum
ETSU032238	Carter	Tennessee		James T Donaldson	ETSU Sceptridium biternatum
ETSU030857	Cocke	Tennessee			ETSU Sceptridium biternatum
ETSU032401	McMinn	Tennessee		James T Donaldson	ETSU Sceptridium biternatum
UOS08641	Franklin	Tennessee	26 Sep 2004	Mary P. Priestley	UOS Sceptridium biternatum
UOS08644	Franklin	Tennessee	12 Nov 1952	E. and D.A. Berkeley	UOS Sceptridium biternatum
BRIT188210	Hancock	Tennessee	05 Sep 1952	Jesse M. Shaver 10928	VDB Botrychium biternatum
BRIT188205	Davidson	Tennessee	28 Sep 1970	R. Kral 41303	VDB Botrychium biternatum
BRIT188220	Sevier	Tennessee	05 Sep 1935	Jesse M. Shaver 2203	VDB Botrychium biternatum
BRIT188209	Hamilton	Tennessee	24 Aug 1994	R. Brown 3	VDB Botrychium biternatum
ETSU030881	Carter	Tennessee			ETSU Sceptridium biternatum
UCHT000269	Bradley	Tennessee	20 Sep 1985	D. Houck 2297	UCHT Sceptridium biternatum
UCHT000261	Hamilton	Tennessee	16 Oct 1985	B. Barrow	UCHT Sceptridium biternatum
UCHT000280	Marion	Tennessee	01 Oct 1981	C. Arellano 12	UCHT Sceptridium biternatum
UCHT006085	Polk	Tennessee			UCHT Sceptridium biternatum
ETSU030875	Cocke	Tennessee			ETSU Sceptridium biternatum
MEM020355	Shelby	Tennessee	11 Nov 2015	S.M. Ballou Jr. 10	MEM Botrychium biternatum
TENN-V-0002047	Hickman	Tennessee	00 August 2003	Dwayne Estes 5248	TENN Botrychium biternatum
MEM012789	Lyon	Kentucky	03 Oct 1977	Raymond Athey 3862	MEM Botrychium biternatum
TENN-V-0002093		Tennessee	21 Sep 1946	R. E. Shanks BB 9	TENN Botrychium biternatum
ETSU031963	Wilson	Tennessee		James T Donaldson	ETSU Sceptridium biternatum
MEM012073	Henderson	Tennessee	17 Aug 1977	Dwight Miller 26	MEM Botrychium biternatum
UCHT000267		Tennessee	07 Sep 1982	K. Thompson, B. Coston	UCHT Sceptridium biternatum
UCHT000281	Marion	Tennessee	1983-Not shown-Not shown	Edwin Bridges and Paul Somers 950	UCHT Sceptridium biternatum
TENN-V-0002099	Polk	Tennessee	07 Sep 1952	Jesse M. Shaver 10903	TENN Botrychium biternatum

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Sceptridium biternatum

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Species Distribution Map

Ecoregions

Synonyms

Specimens and Distribution

Plant Photos